From what I got, it virtualizes everything but CPUs and memory. Each Atom sees virtualized network connections, disk storage and whatever else the custom chipset wants to present. I am not sure if you can make the 8 Atoms on each board see each other as it they were an 8-way SMP box, but I am quite sure you can't make them believe they are a single-image, 512-processor machine.
From what I understood, they'd appear to be separate systems.
The whole point of their custom silicon was to share certain resources between the various CPUs as systems. If it were the equivalent of, say, a 2-CPU motherboard, none of that would have been necessary since they don't each expect their own I/O coprocessors, memory banks, etc.
http://news.ycombinator.com/item?id=1429628