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I think the key differentiator is Prefrontal Synthesis.

https://en.m.wikipedia.org/wiki/Prefrontal_synthesis

This is the ability to mentally compose conceptual objects in complex relationships.

In fact I don’t think there really is one single significant differentiator. There are many. However without this one technology beyond simple single function tools such as a hand axe or pointed stick, and simple linguistic statements wouldn’t be be possible.

The linguistic composition described in the article sound similar if dramatically more primitive, but I don’t think we can assume it comes from a common ancestor. It may well be that this capability has a different neurological basis, since these animals don’t have a prefrontal cortex. So it seems plausible this is a case or parallel evolution of a very rudimentary similar feature. Fascinating stuff though.






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