How is that pdf made interactive? It has options to toggle the behaviour, which work even in an in browser pdf viewer. I did not think PDFs could do that.
PDFs can do a lot more than show static content. There was one time where Adobe strongly advocated for PDF to be the page format of what would come to be called "The World Wide Web". Where we have HTML now, Adobe wanted PDF. Thankfully that did not happen. But I suspect it would have made more sense technically than [whatever this mess is that we have now.]
The microtype user manual shows how much thought has gone into it: https://mirror.foobar.to/CTAN/macros/latex/contrib/microtype...